About the Authors

Francesca Luca

Contributed equally to this work with: Francesca Luca, Joseph C. Maranville

Current address: Center for Molecular Medicine and Genetics, Wayne State University, Detroit, Michigan, United States of America

Affiliation Department of Human Genetics, University of Chicago, Chicago, Illinois, United States of America

Joseph C. Maranville

Contributed equally to this work with: Francesca Luca, Joseph C. Maranville

Affiliation Department of Human Genetics, University of Chicago, Chicago, Illinois, United States of America

Allison L. Richards

Current address: Department of Genetics, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Affiliation Department of Human Genetics, University of Chicago, Chicago, Illinois, United States of America

David B. Witonsky

Affiliation Department of Human Genetics, University of Chicago, Chicago, Illinois, United States of America

Matthew Stephens

Affiliations Department of Human Genetics, University of Chicago, Chicago, Illinois, United States of America, Department of Statistics, University of Chicago, Chicago, Illinois, United States of America

Anna Di Rienzo

dirienzo@bsd.uchicago.edu

Affiliation Department of Human Genetics, University of Chicago, Chicago, Illinois, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: FL JCM AD. Performed the experiments: FL JCM ALR. Analyzed the data: FL JCM DBW. Contributed reagents/materials/analysis tools: MS. Wrote the paper: FL JCM MS AD.