About the Authors

Kunie Matsuoka

matsuoka-kn@igakuken.or.jp

Affiliation Mammalian Genetics Project, Tokyo Metropolitan Institute of Medical Science, Tokyo, Japan

Hiroshi Shitara

Affiliation Laboratory for Transgenic Technology, Tokyo Metropolitan Institute of Medical Science, Tokyo, Japan

Choji Taya

Affiliation Laboratory for Transgenic Technology, Tokyo Metropolitan Institute of Medical Science, Tokyo, Japan

Kenji Kohno

Affiliation Graduate School of Biological Sciences, Nara Institute of Science and Technology, Nara, Japan

Yoshiaki Kikkawa

Affiliation Mammalian Genetics Project, Tokyo Metropolitan Institute of Medical Science, Tokyo, Japan

Hiromichi Yonekawa

Affiliation Laboratory for Transgenic Technology, Tokyo Metropolitan Institute of Medical Science, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KM HY. Performed the experiments: KM HS. Analyzed the data: KM HS HY. Contributed reagents/materials/analysis tools: KM HS CT KK YK. Wrote the paper: KM HY.