About the Authors

Tomohiro Nishimura

Affiliations Faculty of Agriculture, Kochi University, Nankoku, Kochi, Japan, The United Graduate School of Agricultural Sciences, Matsuyama, Ehime University, Ehime, Japan

Shinya Sato

Affiliations Royal Botanic Garden Edinburgh, Edinburgh, United Kingdom, Cardiff University, Cardiff, Wales, United Kingdom

Wittaya Tawong

Affiliations Faculty of Agriculture, Kochi University, Nankoku, Kochi, Japan, The United Graduate School of Agricultural Sciences, Matsuyama, Ehime University, Ehime, Japan

Hiroshi Sakanari

Affiliation Faculty of Agriculture, Kochi University, Nankoku, Kochi, Japan

Keita Uehara

Affiliation Faculty of Agriculture, Kochi University, Nankoku, Kochi, Japan

Md Mahfuzur Rahman Shah

Affiliations Faculty of Science, University of the Ryukyus, Nakagami District, Okinawa, Japan, College of Ocean Science, Jeju National University, Jeju, South Korea

Shoichiro Suda

Affiliation Faculty of Science, University of the Ryukyus, Nakagami District, Okinawa, Japan

Takeshi Yasumoto

Affiliation National Research Institute of Fisheries Science, Yokohama, Kanagawa, Japan

Yohsuke Taira

Affiliation Okinawa Institute of Science and Technology Evolutionary Systems Biology Unit, Kunigami District, Okinawa, Japan

Haruo Yamaguchi

Affiliation Faculty of Agriculture, Kochi University, Nankoku, Kochi, Japan

Masao Adachi

madachi@kochi-u.ac.jp

Affiliation Faculty of Agriculture, Kochi University, Nankoku, Kochi, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TN MMR S. Sato S. Suda MA. Performed the experiments: TN MMR. Analyzed the data: TN S. Sato MA. Contributed reagents/materials/analysis tools: TN S. Sato WT HS KU MMR S. Suda TY YT HY MA. Wrote the paper: TN S. Sato MA.