About the Authors

Anu Lipsanen

Affiliation Institute of Clinical Medicine-Neurology, University of Eastern Finland, Kuopio, Finland

Stefanie Flunkert

Affiliation QPS Austria GmbH, Grambach, Austria

Kristina Kuptsova

Affiliation Institute of Clinical Medicine-Neurology, University of Eastern Finland, Kuopio, Finland

Mikko Hiltunen

Affiliation Institute of Clinical Medicine-Neurology, University of Eastern Finland, Kuopio, Finland

Manfred Windisch

Affiliation QPS Austria GmbH, Grambach, Austria

Birgit Hutter-Paier

Affiliation QPS Austria GmbH, Grambach, Austria

Jukka Jolkkonen

jukka.jolkkonen@uef.fi

Affiliation Institute of Clinical Medicine-Neurology, University of Eastern Finland, Kuopio, Finland

Competing Interests

The following authors affiliated with QPS Austria GmbH (formerly JSW Life Sciences) are employed by the company: Stefanie Flunkert, Manfred Windisch and Birgit Hutter-Paier. All the other authors report no disclosures. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: JJ MH BHP MW. Performed the experiments: AL SF KK. Analyzed the data: AL SF KK. Wrote the paper: JJ MH BHP MW.