About the Authors

Linda D. Stewart

Affiliation Institute for Global Food Security, School of Biological Sciences, Queen's University Belfast, Belfast, Northern Ireland, United Kingdom

James McNair

Affiliation Veterinary Sciences Division, Agri-Food and Biosciences Institute for Northern Ireland, Stormont, Belfast, Northern Ireland, United Kingdom

Lyanne McCallan

Affiliation Veterinary Sciences Division, Agri-Food and Biosciences Institute for Northern Ireland, Stormont, Belfast, Northern Ireland, United Kingdom

Alan Gordon

Affiliation Biometrics Division, Agri-Food and Biosciences Institute for Northern Ireland, Newforge Lane, Belfast, Northern Ireland, United Kingdom

Irene R. Grant

i.grant@qub.ac.uk

Affiliation Institute for Global Food Security, School of Biological Sciences, Queen's University Belfast, Belfast, Northern Ireland, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: IRG JMcN. Performed the experiments: LDS LMcC. Analyzed the data: IRG AG. Wrote the paper: LDS IRG JMcN LMcC.