About the Authors

Shinya Yamamoto


Affiliation Primate Research Institute, Kyoto University, Inuyama, Aichi, Japan

Tatyana Humle

Affiliation School of Anthropology and Conservation, University of Kent, Canterbury, Kent, United Kingdom

Masayuki Tanaka

Affiliation Wildlife Research Center, Kyoto University, Sakyo-ku, Kyoto, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SY TH MT. Performed the experiments: SY. Analyzed the data: SY. Wrote the paper: SY TH MT.