About the Authors

Qi Xu

Contributed equally to this work with: Qi Xu, WenMing Zhao

Affiliation Key Laboratory of Animal Genetics & Breeding and Molecular Design of Jiangsu Province, Yangzhou University, Yangzhou, People's Republic of China

WenMing Zhao

Contributed equally to this work with: Qi Xu, WenMing Zhao

Affiliation Key Laboratory of Animal Genetics & Breeding and Molecular Design of Jiangsu Province, Yangzhou University, Yangzhou, People's Republic of China

Yang Chen

Affiliation Key Laboratory of Animal Genetics & Breeding and Molecular Design of Jiangsu Province, Yangzhou University, Yangzhou, People's Republic of China

YiYu Tong

Affiliation Key Laboratory of Animal Genetics & Breeding and Molecular Design of Jiangsu Province, Yangzhou University, Yangzhou, People's Republic of China

GuangHui Rong

Affiliation Key Laboratory of Animal Genetics & Breeding and Molecular Design of Jiangsu Province, Yangzhou University, Yangzhou, People's Republic of China

ZhengYang Huang

Affiliation Key Laboratory of Animal Genetics & Breeding and Molecular Design of Jiangsu Province, Yangzhou University, Yangzhou, People's Republic of China

Yang Zhang

Affiliation Key Laboratory of Animal Genetics & Breeding and Molecular Design of Jiangsu Province, Yangzhou University, Yangzhou, People's Republic of China

GuoBing Chang

Affiliation Key Laboratory of Animal Genetics & Breeding and Molecular Design of Jiangsu Province, Yangzhou University, Yangzhou, People's Republic of China

XinSheng Wu

Affiliation Key Laboratory of Animal Genetics & Breeding and Molecular Design of Jiangsu Province, Yangzhou University, Yangzhou, People's Republic of China

GuoHong Chen

ghchen@yzu.edu.cn

Affiliation Key Laboratory of Animal Genetics & Breeding and Molecular Design of Jiangsu Province, Yangzhou University, Yangzhou, People's Republic of China

Competing Interests

The authors have the following interests: Jiangsu Lihua Animal Husbandry CO. LTD provided the animal sample. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: GHC WMZ QX. Performed the experiments: QX WMZ. Analyzed the data: QX YC YYT GHR ZYH. Contributed reagents/materials/analysis tools: QX YZ GBC XSW. Wrote the paper: QU WMZ.