About the Authors

Eileen C. Spindler

Current address: OPX Biotechnologies, Inc., Boulder, Colorado, United States of America

Affiliation Department of Biological and Chemical Engineering, University of Colorado Boulder, Boulder, Colorado, United States of America

Nanette R. Boyle

Affiliation Department of Biological and Chemical Engineering, University of Colorado Boulder, Boulder, Colorado, United States of America

Robert E. W. Hancock

Affiliation Centre for Microbial Diseases and Immunity Research, University of British Columbia, Vancouver, Canada

Ryan T. Gill

rtg@colorado.edu

Affiliation Department of Biological and Chemical Engineering, University of Colorado Boulder, Boulder, Colorado, United States of America

Competing Interests

Eileen Spindler is currently employed by OPX Biotechnologies, Inc. which licenses unrelated technologies from the University of Colorado Boulder. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials as Dr. Spindler was a graduate student when this work was completed and had no affiliation with OPX.

Author Contributions

Conceived and designed the experiments: ECS REWH RTG. Performed the experiments: ECS. Analyzed the data: ECS. Wrote the paper: ECS NRB REWH RTG.