About the Authors

Maia M. Chan

Affiliation Department of Comparative Medicine, University of Washington, Seattle, Washington, United States of America

Jason M. Wooden

Affiliation Puget Sound Blood Center, Seattle, Washington, United States of America

Mark Tsang

Affiliation Department of Comparative Medicine, University of Washington, Seattle, Washington, United States of America

Diana M. Gilligan

Affiliation Puget Sound Blood Center, Seattle, Washington, United States of America

Dinesh K. Hirenallur-S

Affiliation Department of Comparative Medicine, University of Washington, Seattle, Washington, United States of America

Greg L. Finney

Affiliation Department of Genome Sciences, University of Washington, Seattle, Washington, United States of America

Eric Rynes

Affiliation Department of Genome Sciences, University of Washington, Seattle, Washington, United States of America

Michael MacCoss

Affiliation Department of Genome Sciences, University of Washington, Seattle, Washington, United States of America

Julita A. Ramirez

Affiliation Department of Comparative Medicine, University of Washington, Seattle, Washington, United States of America

Heon Park

Affiliation Department of Comparative Medicine, University of Washington, Seattle, Washington, United States of America

Brian M. Iritani

biritani@uw.edu

Affiliation Department of Comparative Medicine, University of Washington, Seattle, Washington, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BMI MMC JMW GLF MM DMG DKH-S JAR. Performed the experiments: BMI MMC JMW MT HP JAR. Wrote the paper: BMI MMC JMW DMG DH JAR. Analyzed the data: JMW MT HP GLF MM ER JAR. Contributed reagents/materials/analysis tools: BMI JMW DMG GLF MM ER.