About the Authors

Ya-pin Li

Contributed equally to this work with: Ya-pin Li, Li-qun Fang

Affiliation State Key Laboratory of Pathogens and Biosecurity, Beijing Institute of Microbiology and Epidemiology, Beijing, People’s Republic of China

Li-qun Fang

Contributed equally to this work with: Ya-pin Li, Li-qun Fang

Affiliation State Key Laboratory of Pathogens and Biosecurity, Beijing Institute of Microbiology and Epidemiology, Beijing, People’s Republic of China

Su-qing Gao

Affiliation Institute of Command Automation, PLA University of Science and Technology, Nanjing, People’s Republic of China

Zhen Wang

Affiliation Institute of Command Automation, PLA University of Science and Technology, Nanjing, People’s Republic of China

Hong-wei Gao

Affiliation State Key Laboratory of Pathogens and Biosecurity, Beijing Institute of Microbiology and Epidemiology, Beijing, People’s Republic of China

Peng Liu

Affiliation Institute of Command Automation, PLA University of Science and Technology, Nanjing, People’s Republic of China

Ze-rui Wang

Affiliation Institute of Command Automation, PLA University of Science and Technology, Nanjing, People’s Republic of China

Yan-li Li

Affiliation State Key Laboratory of Pathogens and Biosecurity, Beijing Institute of Microbiology and Epidemiology, Beijing, People’s Republic of China

Xu-guang Zhu

Affiliation Institute of Command Automation, PLA University of Science and Technology, Nanjing, People’s Republic of China

Xin-lou Li

Affiliation State Key Laboratory of Pathogens and Biosecurity, Beijing Institute of Microbiology and Epidemiology, Beijing, People’s Republic of China

Bo Xu

Affiliation Institute of Command Automation, PLA University of Science and Technology, Nanjing, People’s Republic of China

Yin-jun Li

Affiliation State Key Laboratory of Pathogens and Biosecurity, Beijing Institute of Microbiology and Epidemiology, Beijing, People’s Republic of China

Hong Yang

Affiliation State Key Laboratory of Pathogens and Biosecurity, Beijing Institute of Microbiology and Epidemiology, Beijing, People’s Republic of China

Sake J. de Vlas

Affiliation Department of Public Health, Erasmus Medical Center (MC), University Medical Center Rotterdam, Rotterdam, The Netherlands

Tao-xing Shi

caowc@bmi.ac.cn (WCC); shitaoxing@sina.com (TXS)

Affiliation State Key Laboratory of Pathogens and Biosecurity, Beijing Institute of Microbiology and Epidemiology, Beijing, People’s Republic of China

Wu-chun Cao

caowc@bmi.ac.cn (WCC); shitaoxing@sina.com (TXS)

Affiliation State Key Laboratory of Pathogens and Biosecurity, Beijing Institute of Microbiology and Epidemiology, Beijing, People’s Republic of China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Provided development software and workplace: HY. Conceived and designed the experiments: YPL LQF TXS WCC. Performed the experiments: BX YJL. Analyzed the data: YPL LQF SQG ZW HWG. Contributed reagents/materials/analysis tools: PL ZRW YLL XGZ XLL. Wrote the paper: YPL LQF SdV.