About the Authors

Melissa R. Christopherson

Contributed equally to this work with: Melissa R. Christopherson, Garret Suen

Affiliation Department of Bacteriology, University of Wisconsin-Madison, Madison, Wisconsin, United States of America

Garret Suen

Contributed equally to this work with: Melissa R. Christopherson, Garret Suen

Affiliation Department of Bacteriology, University of Wisconsin-Madison, Madison, Wisconsin, United States of America

Shanti Bramhacharya

Affiliation Department of Bacteriology, University of Wisconsin-Madison, Madison, Wisconsin, United States of America

Kelsea A. Jewell

Affiliation Department of Bacteriology, University of Wisconsin-Madison, Madison, Wisconsin, United States of America

Frank O. Aylward

Affiliations Department of Bacteriology, University of Wisconsin-Madison, Madison, Wisconsin, United States of America, Department of Energy, Great Lakes Bioenergy Research Center, University of Wisconsin-Madison, Madison, Wisconsin, United States of America

David Mead

Affiliations Department of Energy, Great Lakes Bioenergy Research Center, University of Wisconsin-Madison, Madison, Wisconsin, United States of America, Lucigen, Middleton, Wisconsin, United States of America

Phillip J. Brumm

pbrumm@c56technologies.com

Affiliations Department of Energy, Great Lakes Bioenergy Research Center, University of Wisconsin-Madison, Madison, Wisconsin, United States of America, C5-6 Technologies, Middleton, Wisconsin, United States of America

Competing Interests

The authors have read the journal’s policy and have the following conflicts: David Mead is employed by Lucigen Corp., a manufacturer of research reagents. Phillip Brumm is employed by C5–6 Technologies Corp., an enzyme discovery company. All work reported here was performed under and supported by subcontract to the GLBRC. No funds from either corporation was used for this research or to support the researchers during performance of this work. The commercial affiliations which the authors have declared do not alter their adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: MRC GS SB KJ FOA DM PB. Performed the experiments: MRC GS SB KJ DM PB. Analyzed the data: MRC GS SB KJ DM PB. Contributed reagents/materials/analysis tools: GS SB KJ FOA DM PB. Wrote the paper: MRC GS PB.