About the Authors

Xiao Hui Xu

Contributed equally to this work with: Xiao Hui Xu, Fang Wang

Current address: Institute of Plant Protection, Shandong Academy of Agricultural Sciences, Ji’nan, Shandong, China

Affiliation State Key Laboratory of Crop Biology, Shandong Key Laboratory of Crop Biology, College of Life Sciences, Shandong Agricultural University, Tai’an, Shandong, China

Fang Wang

Contributed equally to this work with: Xiao Hui Xu, Fang Wang

Affiliation State Key Laboratory of Crop Biology, Shandong Key Laboratory of Crop Biology, College of Life Sciences, Shandong Agricultural University, Tai’an, Shandong, China

Hao Chen

Current address: Forest Biotechnology Group, Department of Forestry and Environmental Resources, North Carolina State University, Raleigh, North Carolina, United States of America

Affiliation State Key Laboratory of Crop Biology, Shandong Key Laboratory of Crop Biology, College of Life Sciences, Shandong Agricultural University, Tai’an, Shandong, China

Wei Sun

Current address: Rice Research Institute, Shandong Academy of Agricultural Sciences, Ji’nan, Shandong, China

Affiliation State Key Laboratory of Crop Biology, Shandong Key Laboratory of Crop Biology, College of Life Sciences, Shandong Agricultural University, Tai’an, Shandong, China

Xian Sheng Zhang

zhangxs@sdau.edu.cn

Affiliation State Key Laboratory of Crop Biology, Shandong Key Laboratory of Crop Biology, College of Life Sciences, Shandong Agricultural University, Tai’an, Shandong, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: XSZ FW. Performed the experiments: XHX HC WS. Analyzed the data: XHX. Wrote the paper: FW XHX XSZ.