About the Authors

Kohei Fujikura

Affiliation Division of Membrane Dynamics, Department of Physiology and Cell Biology, Kobe University Graduate School of Medicine, Kobe, Hyogo, Japan

Tomiyoshi Setsu

Affiliation Division of Developmental Neurobiology, Department of Physiology and Cell Biology, Kobe University Graduate School of Medicine, Kobe, Hyogo, Japan

Kenji Tanigaki

Affiliation Research Institute, Shiga Medical Center, Moriyama, Shiga, Japan

Takaya Abe

Affiliation Laboratory for Animal Resources and Genetic Engineering, RIKEN Center for Developmental Biology, Kobe, Hyogo, Japan

Hiroshi Kiyonari

Affiliation Laboratory for Animal Resources and Genetic Engineering, RIKEN Center for Developmental Biology, Kobe, Hyogo, Japan

Toshio Terashima

Affiliation Division of Developmental Neurobiology, Department of Physiology and Cell Biology, Kobe University Graduate School of Medicine, Kobe, Hyogo, Japan

Toshiaki Sakisaka

sakisaka@ med.kobe-u.ac.jp

Affiliation Division of Membrane Dynamics, Department of Physiology and Cell Biology, Kobe University Graduate School of Medicine, Kobe, Hyogo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TS (Toshiaki Sakisaka) TT. Performed the experiments: KF TS (Toshiaki Sakisaka) KT TA HK TT TS (Tomiyoshi Setsu). Analyzed the data: KF TS (Toshiaki Sakisaka) KT TT TS (Tomiyoshi Setsu). Contributed reagents/materials/analysis tools: TS (Toshiaki Sakisaka) KT TT. Wrote the paper: TS (Toshiaki Sakisaka) KF.