About the Authors

Vishnu Hosur

Contributed equally to this work with: Vishnu Hosur, Melissa L. Cox

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Melissa L. Cox

Contributed equally to this work with: Vishnu Hosur, Melissa L. Cox

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Lisa M. Burzenski

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Rebecca L. Riding

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Lynn Alley

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Bonnie L. Lyons

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Anoop Kavirayani

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Kimberly A. Martin

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Gregory A. Cox

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Kenneth R. Johnson

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Leonard D. Shultz

lenny.shultz@jax.org

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MC VH LS. Performed the experiments: MC VH LB RR LA KM. Analyzed the data: MC VH AK BL GC KJ LS. Contributed reagents/materials/analysis tools: MC VH GC LS. Wrote the paper: MC VH KJ LS.