About the Authors

Daniel Hand

Affiliation School of Biosciences, University of Birmingham, Birmingham, United Kingdom

Corrin Wallis

Affiliation WALTHAM® Centre for Pet Nutrition, Melton Mowbray, Leicestershire, United Kingdom

Alison Colyer

Affiliation WALTHAM® Centre for Pet Nutrition, Melton Mowbray, Leicestershire, United Kingdom

Charles W. Penn

c.w.penn@bham.ac.uk

Affiliation School of Biosciences, University of Birmingham, Birmingham, United Kingdom

Competing Interests

The authors declare a contribution to funding of this project by WALTHAM Centre for Pet Nutrition, to whom CWP is a consultant and by whom DH was supported in part during the study through partial funding of his studentship. This competing interest does not alter authors‚ adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: DH CW CWP. Performed the experiments: DH. Analyzed the data: DH CW AC. Contributed reagents/materials/analysis tools: DH AC CW. Wrote the paper: DH AC CW CWP.