About the Authors

Yuriko Komine

Affiliation Division of Brain Biology, National Institute for Basic Biology, Okazaki, Japan

Keizo Takao

Affiliations Section of Behavior Patterns, National Institute for Physiological Sciences, Okazaki, Japan, Genetic Engineering and Functional Genomics Group, Frontier Technology Center, Kyoto University Graduate School of Medicine, Kyoto, Japan, Division of Systems Medical Science, Institute for Comprehensive Medical Science, Fujita Health University, Toyoake, Japan, Core Research for Evolutional Science and Technology (CREST), Japan Science and Technology Agency (JST), Kawaguchi, Japan

Tsuyoshi Miyakawa

Affiliations Section of Behavior Patterns, National Institute for Physiological Sciences, Okazaki, Japan, Genetic Engineering and Functional Genomics Group, Frontier Technology Center, Kyoto University Graduate School of Medicine, Kyoto, Japan, Division of Systems Medical Science, Institute for Comprehensive Medical Science, Fujita Health University, Toyoake, Japan, Core Research for Evolutional Science and Technology (CREST), Japan Science and Technology Agency (JST), Kawaguchi, Japan

Tetsuo Yamamori

yamamori@nibb.ac.jp

Affiliation Division of Brain Biology, National Institute for Basic Biology, Okazaki, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YK KT TM TY. Performed the experiments: YK KT. Analyzed the data: YK KT. Contributed reagents/materials/analysis tools: YK KT TM. Wrote the paper: YK KT TY. Conceived and designed the test battery experiments: TM.