About the Authors

Brad H. McRae

bmcrae@tnc.org

Affiliation The Nature Conservancy, North America Region, Seattle, Washington, United States of America

Sonia A. Hall

Affiliation The Nature Conservancy, Washington Chapter, Wenatchee, Washington, United States of America

Paul Beier

Affiliation School of Forestry and Merriam-Powell Center for Environmental Research, Northern Arizona University, Flagstaff, Arizona, United States of America

David M. Theobald

Affiliation National Park Service, Inventory and Monitoring Division, Fort Collins, Colorado, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BHM. Performed the experiments: BHM. Analyzed the data: BHM. Contributed reagents/materials/analysis tools: BHM. Wrote the paper: BHM SAH PB DMT. Suggested refinements to methods and analyses: SAH, PB, DMT.