About the Authors

Jennifer R. Bailey

Affiliation School of Veterinary Science, University of Bristol, Bristol, United Kingdom

Paul W. Bland

Contributed equally to this work with: Paul W. Bland, John F. Tarlton

Affiliations School of Veterinary Science, University of Bristol, Bristol, United Kingdom, Mucosal Immunobiology and Vaccine Center (MIVAC), Department of Microbiology and Immunology, University of Gothenburg, Gothenburg, Sweden

John F. Tarlton

Contributed equally to this work with: Paul W. Bland, John F. Tarlton

John.Tarlton@bris.ac.uk

Affiliation School of Veterinary Science, University of Bristol, Bristol, United Kingdom

Iain Peters

Affiliation Molecular Testing, Innovation Centre, University of Exeter, Exeter, United Kingdom

Moganaden Moorghen

Affiliation Histopathology Department, St Mark’s Hospital, Harrow, United Kingdom

Paul A. Sylvester

Affiliation Department of Surgery, Bristol Royal Infirmary, Bristol, United Kingdom

Christopher S. J. Probert

Affiliation Department of Gastroenterology, University of Liverpool, Liverpool, United Kingdom

Christine V. Whiting

Affiliation School of Veterinary Science, University of Bristol, Bristol, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CVW PWB CSJP JFT JRB. Performed the experiments: CVW JRB. Analyzed the data: CVW JRB PWB MM. Contributed reagents/materials/analysis tools: JFT MM PAS IP. Wrote the paper: CVW JRB PWB JFT CSJP.