About the Authors

Shunsuke Yagi

Affiliation Laboratory of Bioimaging and Cell Signaling, Graduate School of Biostudies, Kyoto University, Yoshida Konoe-cho, Sakyo-ku, Kyoto, Japan

Michiyuki Matsuda

Affiliations Laboratory of Bioimaging and Cell Signaling, Graduate School of Biostudies, Kyoto University, Yoshida Konoe-cho, Sakyo-ku, Kyoto, Japan, Department of Pathology and Biology of Diseases, Graduate School of Medicine, Kyoto University, Yoshida Konoe-cho, Sakyo-ku, Kyoto, Japan

Etsuko Kiyokawa

kiyokawa@kanazawa-med.ac.jp

Affiliations Department of Pathology and Biology of Diseases, Graduate School of Medicine, Kyoto University, Yoshida Konoe-cho, Sakyo-ku, Kyoto, Japan, Department of Oncologic Pathology, Kanazawa Medical University, Uchinada, Kahoku-gun, Ishikawa, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SY MM EK. Performed the experiments: SY. Analyzed the data: SY EK. Contributed reagents/materials/analysis tools: SY MM EK. Wrote the paper: SY MM EK.