About the Authors

Than J. Boves

tboves@illinois.edu

Current address: Department of Natural Resources and Environmental Science, University of Illinois, Urbana, Illinois, United States of America

Affiliation Department of Forestry, Wildlife, and Fisheries, University of Tennessee, Knoxville, Tennessee, United States of America

David A. Buehler

Affiliation Department of Forestry, Wildlife, and Fisheries, University of Tennessee, Knoxville, Tennessee, United States of America

James Sheehan

Affiliation West Virginia Cooperative Fish and Wildlife Research Unit, Division of Forestry and Natural Resources, West Virginia University, Morgantown, West Virginia, United States of America

Petra Bohall Wood

Affiliation U.S. Geological Survey, West Virginia Cooperative Fish and Wildlife Research Unit, West Virginia University, Morgantown, West Virginia, United States of America

Amanda D. Rodewald

Affiliation School of Environment and Natural Resources, Ohio State University, Columbus, Ohio, United States of America

Jeffrey L. Larkin

Affiliation Department of Biology, Indiana University of Pennsylvania, Indiana, Pennsylvania, United States of America

Patrick D. Keyser

Affiliation Department of Forestry, Wildlife, and Fisheries, University of Tennessee, Knoxville, Tennessee, United States of America

Felicity L. Newell

Affiliation School of Environment and Natural Resources, Ohio State University, Columbus, Ohio, United States of America

Gregory A. George

Affiliation West Virginia Cooperative Fish and Wildlife Research Unit, Division of Forestry and Natural Resources, West Virginia University, Morgantown, West Virginia, United States of America

Marja H. Bakermans

Affiliation School of Environment and Natural Resources, Ohio State University, Columbus, Ohio, United States of America

Andrea Evans

Affiliation Department of Biology, Indiana University of Pennsylvania, Indiana, Pennsylvania, United States of America

Tiffany A. Beachy

Affiliation Department of Forestry, Wildlife, and Fisheries, University of Tennessee, Knoxville, Tennessee, United States of America

Molly E. McDermott

Affiliation West Virginia Cooperative Fish and Wildlife Research Unit, Division of Forestry and Natural Resources, West Virginia University, Morgantown, West Virginia, United States of America

Kelly A. Perkins

Affiliation West Virginia Cooperative Fish and Wildlife Research Unit, Division of Forestry and Natural Resources, West Virginia University, Morgantown, West Virginia, United States of America

Matthew White

Affiliation Department of Biology, Indiana University of Pennsylvania, Indiana, Pennsylvania, United States of America

T. Bently Wigley

Affiliation National Council for Air and Stream Improvement, Inc., Clemson, South Carolina, United States of America

Competing Interests

MeadWestvaco Corporation provided some funding, however they did not ask to review/comment on our results prior to publishing and their financial contribution had no contingencies with respect to the study. Conservation science is often funded via public-private partnerships such as this. In addition, this does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: TJB DAB JS PBW ADR JLL PDK TAB FLN MHB AE GAG MEM KAP TBW. Performed the experiments: TJB DAB JS PBW ADR JLL TAB FLN MHB AE GAG MEM KAP. Analyzed the data: TJB DAB. Wrote the paper: TJB DAB PDK ADR PBW TBW JLL.