About the Authors

Mark W. Neff

mark.neff@vai.org

Affiliation Laboratory of Neurogenetics and Canine Behavior, Van Andel Research Institute, Grand Rapids, Michigan, United States of America

John S. Beck

Affiliation Laboratory of Neurogenetics and Canine Behavior, Van Andel Research Institute, Grand Rapids, Michigan, United States of America

Julie M. Koeman

Affiliation Laboratory of Neurogenetics and Canine Behavior, Van Andel Research Institute, Grand Rapids, Michigan, United States of America

Elissa Boguslawski

Affiliation Laboratory of Neurogenetics and Canine Behavior, Van Andel Research Institute, Grand Rapids, Michigan, United States of America

Lisa Kefene

Affiliation Laboratory of Neurogenetics and Canine Behavior, Van Andel Research Institute, Grand Rapids, Michigan, United States of America

Andrew Borgman

Affiliation Laboratory of Neurogenetics and Canine Behavior, Van Andel Research Institute, Grand Rapids, Michigan, United States of America

Alison L. Ruhe

Affiliations Laboratory of Neurogenetics and Canine Behavior, Van Andel Research Institute, Grand Rapids, Michigan, United States of America, Program for Canine Health and Performance, Translational Genomics Research Institute, Phoenix, Arizona, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MWN JSB JMK ALR. Analyzed the data: AB. Wrote the paper: MWN. Performed the experiments and collected data: JSB JMK ALR EB LK.