About the Authors

Keiko Miyadera

Current address: Department of Clinical Studies, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Affiliation Department of Veterinary Medicine, University of Cambridge, Cambridge, United Kingdom

Ian Brierley

Affiliation Virology Division, Department of Pathology, University of Cambridge, Cambridge, United Kingdom

Jesús Aguirre-Hernández

Affiliation Department of Veterinary Medicine, University of Cambridge, Cambridge, United Kingdom

Cathryn S. Mellersh

Affiliation Canine Genetics Section, Animal Health Trust, Newmarket, United Kingdom

David R. Sargan

drs20@cam.ac.uk

Affiliation Department of Veterinary Medicine, University of Cambridge, Cambridge, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KM IB CSM DRS. Performed the experiments: KM. Analyzed the data: KM JAH DRS. Contributed reagents/materials/analysis tools: KM IB CSM DRS. Wrote the paper: DRS KM. Critical revision of manuscript: IB JAH CSM.