About the Authors

Smriti Shringi

Affiliation Department of Veterinary Microbiology and Pathology, Washington State University, Pullman, Washington, United States of America

Carrie Schmidt

Affiliation Department of Veterinary Microbiology and Pathology, Washington State University, Pullman, Washington, United States of America

Kaya Katherine

Affiliation Department of Veterinary Microbiology and Pathology, Washington State University, Pullman, Washington, United States of America

Kelly A. Brayton

Affiliation Department of Veterinary Microbiology and Pathology, Washington State University, Pullman, Washington, United States of America

Dale D. Hancock

Affiliation Department of Veterinary Clinical Sciences, Washington State University, Pullman, Washington, United States of America

Thomas E. Besser

tbesser@vetmed.wsu.edu

Affiliation Department of Veterinary Microbiology and Pathology, Washington State University, Pullman, Washington, United States of America

Competing Interests

The authors have read the journal’s policy and have the following conflict. One of the authors, Kelly A. Brayton, is serving on the editorial board of PLOS ONE. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: TEB SS. Performed the experiments: SS CS KK. Analyzed the data: SS TEB KAB DDH. Contributed reagents/materials/analysis tools: TEB. Wrote the paper: TEB SS KAB. Drafted the paper: SS. Revised the paper: SS TEB KAB. Final approval of the paper: SS TEB.