About the Authors

Jin-Sil Park

Contributed equally to this work with: Jin-Sil Park, Mi-Kyung Park

Affiliation The Rheumatism Research Center, Catholic Research Institute of Medical Science, The Catholic University of Korea, Seoul, Republic of Korea

Mi-Kyung Park

Contributed equally to this work with: Jin-Sil Park, Mi-Kyung Park

Affiliation The Rheumatism Research Center, Catholic Research Institute of Medical Science, The Catholic University of Korea, Seoul, Republic of Korea

Hye-Joa Oh

Affiliation The Rheumatism Research Center, Catholic Research Institute of Medical Science, The Catholic University of Korea, Seoul, Republic of Korea

Yun-Ju Woo

Affiliation The Rheumatism Research Center, Catholic Research Institute of Medical Science, The Catholic University of Korea, Seoul, Republic of Korea

Mi-Ae Lim

Affiliation The Rheumatism Research Center, Catholic Research Institute of Medical Science, The Catholic University of Korea, Seoul, Republic of Korea

Jong-Ho Lee

Affiliation Department of Cell and Developmental Biology, School of Dentistry, Seoul National University, Seoul, Republic of Korea

Ji Hyeon Ju

Affiliation The Rheumatism Research Center, Catholic Research Institute of Medical Science, The Catholic University of Korea, Seoul, Republic of Korea

Young Ok Jung

Affiliation Division of Rheumatology, Department of Internal Medicine, Hallym University Kang-Nam Sacred Heart Hospital, Seoul, Republic of Korea

Zang Hee Lee

Affiliation Department of Cell and Developmental Biology, School of Dentistry, Seoul National University, Seoul, Republic of Korea

Sung-Hwan Park

Affiliation The Rheumatism Research Center, Catholic Research Institute of Medical Science, The Catholic University of Korea, Seoul, Republic of Korea

Ho-Youn Kim

Affiliation The Rheumatism Research Center, Catholic Research Institute of Medical Science, The Catholic University of Korea, Seoul, Republic of Korea

Mi-La Cho

rmin6403@hanmail.net (J-KM); iammila@catholic.ac.kr (M-LC)

These authors also contributed equally to this work.

Affiliations The Rheumatism Research Center, Catholic Research Institute of Medical Science, The Catholic University of Korea, Seoul, Republic of Korea, Immune Tolerance Research Center, Convergent Research Consortium for Immunologic Disease (CRCID); The Catholic University of Korea College of Medicine, Seoul, Republic of Korea

Jun-Ki Min

rmin6403@hanmail.net (J-KM); iammila@catholic.ac.kr (M-LC)

These authors also contributed equally to this work.

Affiliation The Rheumatism Research Center, Catholic Research Institute of Medical Science, The Catholic University of Korea, Seoul, Republic of Korea

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MLC JKM JSP MKP. Performed the experiments: JSP MKP HJO YJW MAL JHL JHJ YOJ. Analyzed the data: MLC JKM JSP MKP JHL ZHL. Contributed reagents/materials/analysis tools: SHP HYK. Wrote the paper: MLC JKM JSP.