About the Authors

Joseph S. Johnson

joseph.johnson@uky.edu

Affiliation Department of Forestry, University of Kentucky, Lexington, Kentucky, United States of America

Michael J. Lacki

Affiliation Department of Forestry, University of Kentucky, Lexington, Kentucky, United States of America

Steven C. Thomas

Affiliation United States National Park Service, Mammoth Cave, Kentucky, United States of America

John F. Grider

Affiliation Department of Biology, University of North Carolina, Greensboro, North Carolina, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JSJ MJL SCT. Performed the experiments: JSJ SCT JFG. Analyzed the data: JSJ MJL SCT JFG. Wrote the paper: JSJ.