About the Authors

Ryohei Thomas Nakano

Affiliation Department of Botany, Graduate School of Science, Kyoto University, Kyoto, Japan

Ryo Matsushima

Current address: Institute of Plant Science and Resources, Okayama University, Kurashiki, Japan

Affiliation Department of Botany, Graduate School of Science, Kyoto University, Kyoto, Japan

Atsushi J. Nagano

Current address: Center for Ecological Research, Kyoto University, Otsu, Japan

Affiliation Department of Botany, Graduate School of Science, Kyoto University, Kyoto, Japan

Yoichiro Fukao

Affiliation Graduate School of Biological Sciences, Nara Institute of Science and Technology, Ikoma, Japan

Masayuki Fujiwara

Affiliation Graduate School of Biological Sciences, Nara Institute of Science and Technology, Ikoma, Japan

Maki Kondo

Affiliation Department of Cell Biology, National Institute for Basic Biology, Okazaki, Japan

Mikio Nishimura

Affiliation Department of Cell Biology, National Institute for Basic Biology, Okazaki, Japan

Ikuko Hara-Nishimura

ihnishi@gr.bot.kyoto-u.ac.jp

Affiliation Department of Botany, Graduate School of Science, Kyoto University, Kyoto, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RTN RM MN IHN. Performed the experiments: RTN RM AJN YF MF MK. Analyzed the data: RTN. Wrote the paper: RTN IHN.