About the Authors

Woochang Hwang

Affiliation The Wilmer Eye Institute, Johns Hopkins University School of Medicine, Baltimore, Maryland, United States of America

Laszlo Hackler Jr.

Affiliation The Wilmer Eye Institute, Johns Hopkins University School of Medicine, Baltimore, Maryland, United States of America

George Wu

Affiliation Department of Biostatistics, The Johns Hopkins University Bloomberg School of Public Health, Baltimore, Maryland, United States of America

Hongkai Ji

Affiliation Department of Biostatistics, The Johns Hopkins University Bloomberg School of Public Health, Baltimore, Maryland, United States of America

Donald J. Zack

Affiliations The Wilmer Eye Institute, Johns Hopkins University School of Medicine, Baltimore, Maryland, United States of America, Department of Molecular Biology and Genetics, Johns Hopkins University School of Medicine, Baltimore, Maryland, United States of America, Department of Neuroscience, Johns Hopkins University School of Medicine, Baltimore, Maryland, United States of America, McKusick-Nathans Institute of Genetics, Johns Hopkins University School of Medicine, Baltimore, Maryland, United States of America

Jiang Qian

jiang.qian@jhmi.edu

Affiliation The Wilmer Eye Institute, Johns Hopkins University School of Medicine, Baltimore, Maryland, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: WH JQ. Performed the experiments: WH GW. Analyzed the data: WH JQ. Wrote the paper: WH JQ. Analysis and advisement: LH HJ DZ.