About the Authors

Yoshitaka Nagamine

Affiliations National Institute of Livestock and Grassland Science, Tsukuba, Japan, The Roslin Institute and R(D)SVS, University of Edinburgh, Midlothian, United Kingdom

Ricardo Pong-Wong

Affiliation The Roslin Institute and R(D)SVS, University of Edinburgh, Midlothian, United Kingdom

Pau Navarro

Affiliation MRC Human Genetics Unit, MRC Institute of Genetics and Molecular Medicine, University of Edinburgh, Western General Hospital, Edinburgh, United Kingdom

Veronique Vitart

Affiliation MRC Human Genetics Unit, MRC Institute of Genetics and Molecular Medicine, University of Edinburgh, Western General Hospital, Edinburgh, United Kingdom

Caroline Hayward

Affiliation MRC Human Genetics Unit, MRC Institute of Genetics and Molecular Medicine, University of Edinburgh, Western General Hospital, Edinburgh, United Kingdom

Igor Rudan

Affiliations Centre for Population Health Sciences, University of Edinburgh, Medical School, Edinburgh, United Kingdom, Croatian Centre for Global Health, Faculty of Medicine, University of Split, Split, Croatia

Harry Campbell

Affiliation Centre for Population Health Sciences, University of Edinburgh, Medical School, Edinburgh, United Kingdom

James Wilson

Affiliation Centre for Population Health Sciences, University of Edinburgh, Medical School, Edinburgh, United Kingdom

Sarah Wild

Affiliation Centre for Population Health Sciences, University of Edinburgh, Medical School, Edinburgh, United Kingdom

Andrew A. Hicks

Affiliation Institute of Genetic Medicine, European Academy Bozen/Bolzano (EURAC), Bolzano, Italy. Affiliated Institute of the University of Lübeck, Germany

Peter P. Pramstaller

Affiliations Institute of Genetic Medicine, European Academy Bozen/Bolzano (EURAC), Bolzano, Italy. Affiliated Institute of the University of Lübeck, Germany, Department of Neurology, General Central Hospital, Bolzano, Italy, Department of Neurology, University of Lübeck, Lübeck, Germany

Nicholas Hastie

Affiliation MRC Human Genetics Unit, MRC Institute of Genetics and Molecular Medicine, University of Edinburgh, Western General Hospital, Edinburgh, United Kingdom

Alan F. Wright

Affiliation MRC Human Genetics Unit, MRC Institute of Genetics and Molecular Medicine, University of Edinburgh, Western General Hospital, Edinburgh, United Kingdom

Chris S. Haley

chris.haley@igmm.ed.ac.uk

Affiliations The Roslin Institute and R(D)SVS, University of Edinburgh, Midlothian, United Kingdom, MRC Human Genetics Unit, MRC Institute of Genetics and Molecular Medicine, University of Edinburgh, Western General Hospital, Edinburgh, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YN RP PN CSH. Performed the experiments: YN RP PN CSH. Analyzed the data: YN RP PN CSH. Contributed reagents/materials/analysis tools: VV CH IR HC JW SW AH PP NH AW. Wrote the paper: YN RP PN CSH VV.