About the Authors

Kshitij Gupta

Affiliation Department of Pathology, School of Dental Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Hariharan Subramanian

Affiliation Department of Pathology, School of Dental Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Andreas Klos

Affiliation Institute of Medical Microbiology and Hospital Epidemiology, Medical School Hannover, Hannover, Germany

Hydar Ali

alih@upenn.edu

Affiliation Department of Pathology, School of Dental Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Competing Interests

AK, a co-author of this manuscript, is an Academic Editor of PLOS ONE. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: HA KG. Performed the experiments: KG HS. Analyzed the data: KG HA. Contributed reagents/materials/analysis tools: AK KG HA. Wrote the paper: KG HA.