About the Authors

Ryuji J. Machida


Affiliation Biodiversity Research Center, Academia Sinica, Taipei, Taiwan

Nancy Knowlton

Affiliation National Museum of Natural History, Smithsonian Institution, Washington, D.C., United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RJM NK. Performed the experiments: RJM. Analyzed the data: RJM. Contributed reagents/materials/analysis tools: NK. Wrote the paper: RJM NK.