About the Authors

Elaine S. Leung

elaineswleung@gmail.com

Affiliations Department of Zoology, University of Otago, Dunedin, New Zealand, School of Surveying, University of Otago, Dunedin, New Zealand

B. Louise Chilvers

Affiliation Department of Conservation, Aquatic & Threats Unit, Wellington, New Zealand

Shinichi Nakagawa

Affiliation Department of Zoology, University of Otago, Dunedin, New Zealand

Antoni B. Moore

Affiliation School of Surveying, University of Otago, Dunedin, New Zealand

Bruce C. Robertson

Affiliation Department of Zoology, University of Otago, Dunedin, New Zealand

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BC. Performed the experiments: BC EL. Analyzed the data: EL. Contributed reagents/materials/analysis tools: EL BC SN AM BR. Wrote the paper: EL. Contributed to conception of manuscript: EL BC SN AM BR. Reviewed and edited manuscript: BC SN AM BR. Provided advice on data and statistical analysis: SN AM BR. Obtained funding and permit: BC.