About the Authors

Marietjie Landman

Marietjie.landman@nmmu.ac.za

Affiliation Centre for African Conservation Ecology, Department of Zoology, Nelson Mandela Metropolitan University, Port Elizabeth, South Africa

David S. Schoeman

Affiliations Centre for African Conservation Ecology, Department of Zoology, Nelson Mandela Metropolitan University, Port Elizabeth, South Africa, Faculty of Science, Health, Education and Engineering, University of the Sunshine Coast, Maroochydore, DC, Queensland, Australia

Anthony J. Hall-Martin

Affiliation Hall-Martin Consulting CC, Somerset West, South Africa

Graham I. H. Kerley

Affiliation Centre for African Conservation Ecology, Department of Zoology, Nelson Mandela Metropolitan University, Port Elizabeth, South Africa

Competing Interests

The International Bank for Reconstruction and Development provided financial support. Anthony J. Hall-Martin is an employee of Hall-Martin Consulting. This does not alter the authors’ adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: ML AJH GIHK. Performed the experiments: ML AJH. Analyzed the data: ML DS. Wrote the paper: ML DS GIHK.