About the Authors

Emily K. Forbes

ekforbes@hotmail.com

Affiliation The Jenner Institute, University of Oxford, Oxford, United Kingdom

Simone C. de Cassan

Affiliation The Jenner Institute, University of Oxford, Oxford, United Kingdom

David Llewellyn

Affiliation The Jenner Institute, University of Oxford, Oxford, United Kingdom

Sumi Biswas

Affiliation The Jenner Institute, University of Oxford, Oxford, United Kingdom

Anna L. Goodman

Affiliation The Jenner Institute, University of Oxford, Oxford, United Kingdom

Matthew G. Cottingham

Affiliation The Jenner Institute, University of Oxford, Oxford, United Kingdom

Carole A. Long

Affiliation Laboratory of Malaria and Vector Research, National Institute of Allergy and Infectious Diseases/National Institutes of Health, Rockville, Maryland, United States of America

Richard J. Pleass

Affiliation Liverpool School of Tropical Medicine, Liverpool, United Kingdom

Adrian V. S. Hill

Affiliation The Jenner Institute, University of Oxford, Oxford, United Kingdom

Fergal Hill

Affiliation Imaxio SA, Lyon, France

Simon J. Draper

Affiliation The Jenner Institute, University of Oxford, Oxford, United Kingdom

Competing Interests

The authors have read the journal’s policy and have the following conflicts: ALG, AVSH, FH and SJD are named inventors on US 12/595 574 and UK PCT/GB2008/01262, US 12/595 351 and UK PCT/GB2008/01271 patent applications covering malaria vectored vaccines and immunization regimes. FH is an employee of Imaxio SA which owns rights to and is developing the C4 bp adjuvant technology. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: EKF AVSH FH SJD. Performed the experiments: EKF SdC DL CAL SJD. Analyzed the data: EKF SdC MGC CAL SJD. Contributed reagents/materials/analysis tools: SB ALG RJP FH. Wrote the paper: EKF SJD.