About the Authors

Jason G. Estes

jasongestes@gmail.com

Affiliations Department of Environmental Studies, Antioch University New England, Keene, New Hampshire, United States of America, Department of Environmental Conservation, University of Massachusetts, Amherst, Massachusetts, United States of America

Nurzhafarina Othman

Affiliations Organisms and Environment Division, School of Biosciences, Cardiff University, Cardiff, United Kingdom, Danau Girang Field Centre, Sabah Wildlife Department, Kota Kinabalu, Sabah, Malaysia

Sulaiman Ismail

Affiliation Hutan, Kinabatangan Orang-utan Conservation Program, Kota Kinabalu, Sabah, Malaysia

Marc Ancrenaz

Affiliations Hutan, Kinabatangan Orang-utan Conservation Program, Kota Kinabalu, Sabah, Malaysia, Sabah Wildlife Department, Kota Kinabalu, Sabah, Malaysia, North of England Zoological Society, Honorary Conservation Fellow, Chester, United Kingdom

Benoit Goossens

Affiliations Organisms and Environment Division, School of Biosciences, Cardiff University, Cardiff, United Kingdom, Danau Girang Field Centre, Sabah Wildlife Department, Kota Kinabalu, Sabah, Malaysia, Sabah Wildlife Department, Kota Kinabalu, Sabah, Malaysia

Laurentius N. Ambu

Affiliation Sabah Wildlife Department, Kota Kinabalu, Sabah, Malaysia

Anna B. Estes

Affiliation Center for Regional Environmental Studies, Dept. of Environmental Sciences, University of Virginia, Charlottesville, North Carolina, United States of America

Peter A. Palmiotto

Affiliation Department of Environmental Studies, Antioch University New England, Keene, New Hampshire, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JGE PAP MA BG LA NO SI. Performed the experiments: JGE SI NO MA BG. Analyzed the data: JGE. Wrote the paper: JGE MA BG PAP ABE.