About the Authors

Indra Adrianto

Affiliation Arthritis and Clinical Immunology Research Program, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, United States of America

Chee Paul Lin

Affiliation Arthritis and Clinical Immunology Research Program, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, United States of America

Jessica J. Hale

Affiliation Arthritis and Clinical Immunology Research Program, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, United States of America

Albert M. Levin

Affiliation Department of Public Health Sciences, Henry Ford Health System, Detroit, Michigan, United States of America

Indrani Datta

Affiliation Department of Public Health Sciences, Henry Ford Health System, Detroit, Michigan, United States of America

Ryan Parker

Affiliation Arthritis and Clinical Immunology Research Program, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, United States of America

Adam Adler

Affiliation Arthritis and Clinical Immunology Research Program, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, United States of America

Jennifer A. Kelly

Affiliation Arthritis and Clinical Immunology Research Program, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, United States of America

Kenneth M. Kaufman

Affiliations Division of Rheumatology, Cincinnati Children’s Hospital Medical Center, Cincinnati, Ohio, United States of America, The United States Department of Veterans Affairs Medical Center, Cincinnati, Ohio, United States of America

Christopher J. Lessard

Affiliations Arthritis and Clinical Immunology Research Program, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, United States of America, Department of Pathology, University of Oklahoma Health Sciences Center, Oklahoma City, Oklahoma, United States of America

Kathy L. Moser

Affiliations Arthritis and Clinical Immunology Research Program, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, United States of America, Department of Pathology, University of Oklahoma Health Sciences Center, Oklahoma City, Oklahoma, United States of America

Robert P. Kimberly

Affiliation Department of Medicine, University of Alabama at Birmingham, Birmingham, Alabama, United States of America

John B. Harley

Affiliations Division of Rheumatology, Cincinnati Children’s Hospital Medical Center, Cincinnati, Ohio, United States of America, The United States Department of Veterans Affairs Medical Center, Cincinnati, Ohio, United States of America

Michael C. Iannuzzi

Affiliation Department of Medicine, SUNY Upstate Medical University, Syracuse, New York, United States of America

Benjamin A. Rybicki

Affiliation Department of Public Health Sciences, Henry Ford Health System, Detroit, Michigan, United States of America

Courtney G. Montgomery

Courtney-Montgomery@omrf.org

Affiliation Arthritis and Clinical Immunology Research Program, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, United States of America

Competing Interests

CGM is a PLoS ONE Editorial Board member. This does not alter the authors’ adherence to all the PLoS ONE policies on sharing data and materials. The authors declare no competing financial interests.

Author Contributions

Conceived and designed the experiments: IA MCI BAR CGM. Performed the experiments: IA CPL AA KMK. Analyzed the data: IA CPL JJH AML ID RP JAK CJL MCI BAR CGM. Contributed reagents/materials/analysis tools: MCI BAR KLM RPK JBH CGM. Wrote the paper: IA CPL JAK CJL MCI BAR CGM. Approved the final draft: IA CPL JJH AML ID RP AA JAK KMK CJL KLM RPK JBH MCI BAR CGM.