About the Authors

Jing Hu

Contributed equally to this work with: Jing Hu, Hongjie Yao

Affiliation Department of Biochemistry and Molecular Biology, Center for Eukaryotic Gene Regulation, Pennsylvania State University, University Park, Pennsylvania, United States of America

Hongjie Yao

Contributed equally to this work with: Jing Hu, Hongjie Yao

Affiliation Department of Biochemistry and Molecular Biology, Center for Eukaryotic Gene Regulation, Pennsylvania State University, University Park, Pennsylvania, United States of America

Fei Gan

Affiliation Department of Biochemistry and Molecular Biology, Center for Eukaryotic Gene Regulation, Pennsylvania State University, University Park, Pennsylvania, United States of America

Anthony Tokarski

Affiliation Department of Biochemistry and Molecular Biology, Center for Eukaryotic Gene Regulation, Pennsylvania State University, University Park, Pennsylvania, United States of America

Yanming Wang

yuw12@psu.edu

Affiliation Department of Biochemistry and Molecular Biology, Center for Eukaryotic Gene Regulation, Pennsylvania State University, University Park, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YW JH HY. Performed the experiments: JH HY FG AT. Analyzed the data: JH HY YW. Wrote the paper: JH HY YW.