About the Authors

Tracey McDole

tsmcdole@yahoo.com

Affiliation Biology Department, San Diego State University, San Diego, California, United States of America

James Nulton

Affiliation Department of Math and Computer Sciences, San Diego State University, San Diego, California, United States of America

Katie L. Barott

Affiliation Biology Department, San Diego State University, San Diego, California, United States of America

Ben Felts

Affiliation Department of Math and Computer Sciences, San Diego State University, San Diego, California, United States of America

Carol Hand

Affiliation Department of Math and Computer Sciences, San Diego State University, San Diego, California, United States of America

Mark Hatay

Affiliation Biology Department, San Diego State University, San Diego, California, United States of America

Hochul Lee

Affiliation Department of Math and Computer Sciences, San Diego State University, San Diego, California, United States of America

Marc O. Nadon

Affiliations Rosenstiel School of Marine and Atmospheric Science, University of Miami, Miami, Florida, United States of America, Joint Institute for Marine and Atmospheric Research, University of Hawai'i at Manoa, Honolulu, Hawaii, United States of America

Bahador Nosrat

Affiliation Biology Department, San Diego State University, San Diego, California, United States of America

Peter Salamon

Affiliation Department of Math and Computer Sciences, San Diego State University, San Diego, California, United States of America

Barbara Bailey

Affiliation Department of Math and Computer Sciences, San Diego State University, San Diego, California, United States of America

Stuart A. Sandin

Affiliation Scripps Institution of Oceanography, University of California San Diego, La Jolla, California, United States of America

Bernardo Vargas-Angel

Affiliation Pacific Islands Fisheries Science Center, NOAA Fisheries, Honolulu, Hawaii, United States of America

Merry Youle

Affiliation Biology Department, San Diego State University, San Diego, California, United States of America

Brian J. Zgliczynski

Affiliation Scripps Institution of Oceanography, University of California San Diego, La Jolla, California, United States of America

Russell E. Brainard

Affiliation Pacific Islands Fisheries Science Center, NOAA Fisheries, Honolulu, Hawaii, United States of America

Forest Rohwer

Affiliation Biology Department, San Diego State University, San Diego, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: FR TM MH SS. Performed the experiments: TM. Analyzed the data: TM MN BN BB. Contributed reagents/materials/analysis tools: RB. Wrote the paper: TM FR MY. Collected microbial data: TM KB MH. Collected and supplied benthic data: BVA. Collected and supplied fish data: BZ. Oversaw metabolic calculations: JN BF CH HL PS.