About the Authors

Stewart T. G. Burgess

stewart.burgess@moredun.ac.uk

Affiliation Moredun Research Institute, Pentlands Science Park, Bush Loan, Edinburgh, Midlothian, Scotland, United Kingdom

Andrew Greer

Affiliation Faculty of Agriculture and Life Sciences, Lincoln University, Lincoln, Canterbury, New Zealand

David Frew

Affiliation Moredun Research Institute, Pentlands Science Park, Bush Loan, Edinburgh, Midlothian, Scotland, United Kingdom

Beth Wells

Affiliation Moredun Research Institute, Pentlands Science Park, Bush Loan, Edinburgh, Midlothian, Scotland, United Kingdom

Edward J. Marr

Affiliation Moredun Research Institute, Pentlands Science Park, Bush Loan, Edinburgh, Midlothian, Scotland, United Kingdom

Alasdair J. Nisbet

Affiliation Moredun Research Institute, Pentlands Science Park, Bush Loan, Edinburgh, Midlothian, Scotland, United Kingdom

John F. Huntley

Affiliation Moredun Research Institute, Pentlands Science Park, Bush Loan, Edinburgh, Midlothian, Scotland, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: STGB AJN JFH. Performed the experiments: STGB AG DF BW EJM. Analyzed the data: STGB EJM BW. Contributed reagents/materials/analysis tools: AG. Wrote the paper: STGB AJN JFH.