About the Authors

Merryl Gelling

merryl.gelling@zoo.ox.ac.uk

Affiliation Wildlife Conservation Research Unit, Department of Zoology, University of Oxford, Recanati-Kaplan Centre, Tubney, United Kingdom

Paul J. Johnson

Affiliation Wildlife Conservation Research Unit, Department of Zoology, University of Oxford, Recanati-Kaplan Centre, Tubney, United Kingdom

Tom P. Moorhouse

Affiliation Wildlife Conservation Research Unit, Department of Zoology, University of Oxford, Recanati-Kaplan Centre, Tubney, United Kingdom

David W. Macdonald

Affiliation Wildlife Conservation Research Unit, Department of Zoology, University of Oxford, Recanati-Kaplan Centre, Tubney, United Kingdom

Competing Interests

D. Gow Consultancies assisted with limited funding towards the larger project. This does not alter the authors’ adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: MG. Performed the experiments: MG TPM. Analyzed the data: MG PJJ. Contributed reagents/materials/analysis tools: MG DWM. Wrote the paper: MG PJJ TPM DWM.