About the Authors

Ailong Wang

Contributed equally to this work with: Ailong Wang, Youqiang Xu

Affiliation State Key Laboratory of Microbial Technology, Shandong University, Jinan, People's Republic of China

Youqiang Xu

Contributed equally to this work with: Ailong Wang, Youqiang Xu

Affiliation State Key Laboratory of Microbial Technology, Shandong University, Jinan, People's Republic of China

Cuiqing Ma

macq@sdu.edu.cn

Affiliation State Key Laboratory of Microbial Technology, Shandong University, Jinan, People's Republic of China

Chao Gao

Affiliation State Key Laboratory of Microbial Technology, Shandong University, Jinan, People's Republic of China

Lixiang Li

Affiliation State Key Laboratory of Microbial Technology, Shandong University, Jinan, People's Republic of China

Yu Wang

Affiliation State Key Laboratory of Microbial Technology, Shandong University, Jinan, People's Republic of China

Fei Tao

Affiliation School of Life Sciences and Biotechnology, Shanghai Jiao Tong University, Shanghai, People's Republic of China

Ping Xu

Affiliations State Key Laboratory of Microbial Technology, Shandong University, Jinan, People's Republic of China, School of Life Sciences and Biotechnology, Shanghai Jiao Tong University, Shanghai, People's Republic of China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: PX CM AW. Performed the experiments: AW YX. Analyzed the data: CM AW YX LL YW. Contributed reagents/materials/analysis tools: CM CG FT PX. Wrote the paper: PX AW YX CM.