About the Authors

Zhen Yao

Affiliation Departments of Medicine and Physiology, University of California San Francisco, San Francisco, California, United States of America

Wan Namkung

Affiliations Departments of Medicine and Physiology, University of California San Francisco, San Francisco, California, United States of America, Yonsei University, College of Pharmacy, Yonsei Institute of Pharmaceutical Sciences, Incheon, Korea

Eun A. Ko

Affiliation Departments of Medicine and Physiology, University of California San Francisco, San Francisco, California, United States of America

Jinhong Park

Affiliation Yonsei University, College of Pharmacy, Yonsei Institute of Pharmaceutical Sciences, Incheon, Korea

Lukmanee Tradtrantip

Affiliation Departments of Medicine and Physiology, University of California San Francisco, San Francisco, California, United States of America

A. S. Verkman

Alan.Verkman@ucsf.edu

Affiliation Departments of Medicine and Physiology, University of California San Francisco, San Francisco, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ASV. Performed the experiments: ZY WN EK JP. Analyzed the data: ZY WN EK JP. Contributed reagents/materials/analysis tools: ZY WN EK JP. Wrote the paper: ASV ZY WN LT.