About the Authors

Junichi Fujinuma

Affiliation Graduate School of Environmental Science, Hokkaido University, Sapporo, Hokkaido, Japan

Rhett D. Harrison


Affiliation Key Laboratory for Tropical Forest Ecology, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Science, Mengla, Yunnan, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JF RDH. Performed the experiments: JF. Analyzed the data: JF. Wrote the paper: JF RDH.