About the Authors

Yang Wang

Contributed equally to this work with: Yang Wang, Congming Wu

Affiliation Key Laboratory of Development and Evaluation of Chemical and Herbal Drugs for Animal Use, College of Veterinary Medicine, China Agricultural University, Beijing, People's Republic of China

Congming Wu

Contributed equally to this work with: Yang Wang, Congming Wu

Affiliation Key Laboratory of Development and Evaluation of Chemical and Herbal Drugs for Animal Use, College of Veterinary Medicine, China Agricultural University, Beijing, People's Republic of China

Qijing Zhang

Affiliation Department of Veterinary Microbiology and Preventive Medicine, College of Veterinary Medicine, Iowa State University, Ames, Iowa, United States of America

Jing Qi

Affiliation Institute of Animal Science and Veterinary Medicine, Shandong Academy of Agricultural Sciences, Jinan, People's Republic of China

Hebing Liu

Affiliation Key Laboratory of Development and Evaluation of Chemical and Herbal Drugs for Animal Use, College of Veterinary Medicine, China Agricultural University, Beijing, People's Republic of China

Yu Wang

Affiliation Key Laboratory of Development and Evaluation of Chemical and Herbal Drugs for Animal Use, College of Veterinary Medicine, China Agricultural University, Beijing, People's Republic of China

Tao He

Affiliation Key Laboratory of Development and Evaluation of Chemical and Herbal Drugs for Animal Use, College of Veterinary Medicine, China Agricultural University, Beijing, People's Republic of China

Licai Ma

Affiliation Key Laboratory of Development and Evaluation of Chemical and Herbal Drugs for Animal Use, College of Veterinary Medicine, China Agricultural University, Beijing, People's Republic of China

Jing Lai

Affiliation Key Laboratory of Development and Evaluation of Chemical and Herbal Drugs for Animal Use, College of Veterinary Medicine, China Agricultural University, Beijing, People's Republic of China

Zhangqi Shen

Affiliation Department of Veterinary Microbiology and Preventive Medicine, College of Veterinary Medicine, Iowa State University, Ames, Iowa, United States of America

Yuqing Liu

Affiliation Institute of Animal Science and Veterinary Medicine, Shandong Academy of Agricultural Sciences, Jinan, People's Republic of China

Jianzhong Shen

sjz@cau.edu.cn

Affiliation Key Laboratory of Development and Evaluation of Chemical and Herbal Drugs for Animal Use, College of Veterinary Medicine, China Agricultural University, Beijing, People's Republic of China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: Yang Wang CW QZ JS. Performed the experiments: Yu Wang JQ HL TH LM JL. Analyzed the data: CW ZS YL. Contributed reagents/materials/analysis tools: HL TH LM JL. Wrote the paper: Yang Wang QZ JS.