About the Authors

Paul J. Wichgers Schreur

Affiliations Central Veterinary Institute, Wageningen UR, Lelystad, The Netherlands, Department of Infectious Diseases and Immunology, Utrecht University, Utrecht, The Netherlands

Christian van Weeghel

Affiliation Central Veterinary Institute, Wageningen UR, Lelystad, The Netherlands

Johanna M. J. Rebel

Affiliation Central Veterinary Institute, Wageningen UR, Lelystad, The Netherlands

Mari A. Smits

Affiliations Central Veterinary Institute, Wageningen UR, Lelystad, The Netherlands, Wageningen Livestock Research, Wageningen UR, Lelystad, The Netherlands

Jos P. M. van Putten

Affiliation Department of Infectious Diseases and Immunology, Utrecht University, Utrecht, The Netherlands

Hilde E. Smith

hilde.smith@wur.nl

Affiliation Central Veterinary Institute, Wageningen UR, Lelystad, The Netherlands

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: PJWS CvW JMJR MAS JPMvP HES. Performed the experiments: PJWS CvW. Analyzed the data: PJWS. Contributed reagents/materials/analysis tools: PJWS CvW JMJR MAS JPMvP HES. Wrote the paper: PJWS JMJR MAS JPMvP HES.