About the Authors

Rachel E. Johnson

Affiliation Division of Virology, National Institute for Biological Standards and Control, Health Protection Agency, Blanche Lane, Potters Bar, Hertfordshire, United Kingdom

Michelle Hamill

Affiliation BioStatistics, National Institute for Biological Standards and Control, Health Protection Agency, Blanche Lane, Potters Bar, Hertfordshire, United Kingdom

Ruth Harvey

Affiliation Division of Virology, National Institute for Biological Standards and Control, Health Protection Agency, Blanche Lane, Potters Bar, Hertfordshire, United Kingdom

Carolyn Nicolson

Affiliation Division of Virology, National Institute for Biological Standards and Control, Health Protection Agency, Blanche Lane, Potters Bar, Hertfordshire, United Kingdom

James S. Robertson

Affiliation Division of Virology, National Institute for Biological Standards and Control, Health Protection Agency, Blanche Lane, Potters Bar, Hertfordshire, United Kingdom

Othmar G. Engelhardt

othmar.engelhardt@nibsc.hpa.org.uk

Affiliation Division of Virology, National Institute for Biological Standards and Control, Health Protection Agency, Blanche Lane, Potters Bar, Hertfordshire, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: REJ JSR OGE. Performed the experiments: REJ RH CN. Analyzed the data: REJ MH RH JSR OGE. Contributed reagents/materials/analysis tools: REJ. Wrote the paper: REJ JSR OGE.