About the Authors

Teruhiko Takahara

takahara@hiroshima-u.ac.jp

Affiliations Research Institute for Humanity and Nature, Kyoto, Japan, Institute for Sustainable Sciences and Development, Hiroshima University, Higashi-Hiroshima, Japan

Toshifumi Minamoto

Affiliation Research Institute for Humanity and Nature, Kyoto, Japan

Hiroki Yamanaka

Affiliation Department of Environmental Solution Technology, Faculty of Science and Technology, Ryukoku University, Otsu, Shiga, Japan

Hideyuki Doi

Affiliation Institute for Sustainable Sciences and Development, Hiroshima University, Higashi-Hiroshima, Japan

Zen'ichiro Kawabata

Affiliation Research Institute for Humanity and Nature, Kyoto, Japan

Competing Interests

The authors have read the journal's policy. Dr. Y. Iseri is working for a private company (the West Japan Engineering Consultants, Inc.). The RIHN possessed all the data and materials, because the RIHN and his company were making a contract of research support. Thus, this does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: TT TM ZK. Performed the experiments: TT TM HY. Analyzed the data: TT TM HY HD. Wrote the paper: TT TM HD.