About the Authors

Shiaw-Wei Tyan

Affiliation Genomics Research Center, Academia Sinica, Taipei, Taiwan

Chih-Hung Hsu

Affiliation Genomics Research Center, Academia Sinica, Taipei, Taiwan

Kai-Lin Peng

Affiliations Genomics Research Center, Academia Sinica, Taipei, Taiwan, Institute of Biochemistry and Molecular Biology, National Yang-Ming University, Taipei, Taiwan

Chun-Chin Chen

Affiliation Genomics Research Center, Academia Sinica, Taipei, Taiwan

Wen-Hung Kuo

Affiliation Department of Surgery, National Taiwan University Hospital, Taipei, Taiwan

Eva Y.-H. P. Lee

Affiliations Genomics Research Center, Academia Sinica, Taipei, Taiwan, Department of Developmental and Cell Biology, University of California Irvine, Irvine, California, United States of America, Department of Biological Chemistry, University of California Irvine, Irvine, California, United States of America

Jin-Yuh Shew

Affiliation Genomics Research Center, Academia Sinica, Taipei, Taiwan

King-Jen Chang

kingjen@ntu.edu.tw (KJC); ljjuan@gate.sinica.edu.tw (LJJ); whlee@uci.edu (WHL)

Affiliations Department of Surgery, National Taiwan University Hospital, Taipei, Taiwan, Cheng Ching General Hospital, Taichung, Taiwan

Li-Jung Juan

kingjen@ntu.edu.tw (KJC); ljjuan@gate.sinica.edu.tw (LJJ); whlee@uci.edu (WHL)

Affiliation Genomics Research Center, Academia Sinica, Taipei, Taiwan

Wen-Hwa Lee

kingjen@ntu.edu.tw (KJC); ljjuan@gate.sinica.edu.tw (LJJ); whlee@uci.edu (WHL)

Affiliations Genomics Research Center, Academia Sinica, Taipei, Taiwan, Department of Biological Chemistry, University of California Irvine, Irvine, California, United States of America

Competing Interests

Based on UCI policy, WHL declares that he serves as member of board of directors of GeneTex Inc. This arrangement has been reviewed and approved by the COI committee of UCI. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: SWT CHH EYHPL JYS LJJ WHL. Performed the experiments: SWT CHH KLP CCC. Analyzed the data: SWT CHH KLP CCC. Contributed reagents/materials/analysis tools: WHK KJC. Wrote the paper: SWT CHH EYHPL LJJ WHL.