About the Authors

Neil Charles Wallace Mackenzie

Affiliation The Roslin Institute and Royal (Dick) School of Veterinary Studies, The University of Edinburgh, Midlothian, Scotland, United Kingdom

Dongxing Zhu

Affiliation The Roslin Institute and Royal (Dick) School of Veterinary Studies, The University of Edinburgh, Midlothian, Scotland, United Kingdom

Elspeth M. Milne

Affiliation The Roslin Institute and Royal (Dick) School of Veterinary Studies, The University of Edinburgh, Midlothian, Scotland, United Kingdom

Rob van 't Hof

Affiliation Rheumatic Diseases Unit, Molecular Medicine Centre, Institute of Genetics and Molecular Medicine, University of Edinburgh, Edinburgh, United Kingdom

Aline Martin

Affiliation University of Tennessee Health Science Center, Memphis, Tennessee, United States of America

Darryl Leigh Quarles

Affiliation University of Tennessee Health Science Center, Memphis, Tennessee, United States of America

José Luis Millán

Affiliation Sanford Children's Health Research Center, Sanford-Burnham Medical Research Institute, La Jolla, California, United States of America

Colin Farquharson

Affiliation The Roslin Institute and Royal (Dick) School of Veterinary Studies, The University of Edinburgh, Midlothian, Scotland, United Kingdom

Vicky Elisabeth MacRae

vicky.macrae@roslin.ed.ac.uk

Affiliation The Roslin Institute and Royal (Dick) School of Veterinary Studies, The University of Edinburgh, Midlothian, Scotland, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: VEM CF JLM DLQ NCWM. Performed the experiments: NCWM DZ VEM AM EMM. Analyzed the data: NCWM DZ VEM RvH EMM. Contributed reagents/materials/analysis tools: NCWM DZ VEM JLM RvH EMM DLQ AM. Wrote the paper: NCWM VEM CF JLM DLQ.