About the Authors

Yucheng Zhang

Contributed equally to this work with: Yucheng Zhang, Linyong Mao

Affiliations College of Horticulture, Northwest A&F University, Key Laboratory of Horticultural Plant Biology and Germplasm Innovation in Northwest China, Ministry of Agriculture, Yangling, Shaanxi, People's Republic of China, College of Enology, Northwest A&F University, Shaanxi Engineering Research Center for Viti-Viniculture, Yangling, Shaanxi, People's Republic of China

Linyong Mao

Contributed equally to this work with: Yucheng Zhang, Linyong Mao

Affiliation Boyce Thompson Institute for Plant Research, Cornell University, Ithaca, New York, United States of America

Hua Wang

Affiliation College of Enology, Northwest A&F University, Shaanxi Engineering Research Center for Viti-Viniculture, Yangling, Shaanxi, People's Republic of China

Chad Brocker

Affiliation Molecular Toxicology and Environmental Health Sciences Program, Department of Pharmaceutical Sciences, University of Colorado Anschutz Medical Center, Aurora, Colorado, United States of America

Xiangjing Yin

Affiliation College of Horticulture, Northwest A&F University, Key Laboratory of Horticultural Plant Biology and Germplasm Innovation in Northwest China, Ministry of Agriculture, Yangling, Shaanxi, People's Republic of China

Vasilis Vasiliou

Affiliation Molecular Toxicology and Environmental Health Sciences Program, Department of Pharmaceutical Sciences, University of Colorado Anschutz Medical Center, Aurora, Colorado, United States of America

Zhangjun Fei

zf25@cornell.edu (ZF); wangxiping@nwsuaf.edu.cn (XW)

Affiliations Boyce Thompson Institute for Plant Research, Cornell University, Ithaca, New York, United States of America, USDA Robert W. Holley Center for Agriculture and Health, Ithaca, New York, United States of America

Xiping Wang

zf25@cornell.edu (ZF); wangxiping@nwsuaf.edu.cn (XW)

Affiliations College of Horticulture, Northwest A&F University, Key Laboratory of Horticultural Plant Biology and Germplasm Innovation in Northwest China, Ministry of Agriculture, Yangling, Shaanxi, People's Republic of China, State Key Laboratory of Crop Stress Biology in Arid Areas, Northwest A&F University, Yangling, Shaanxi, People's Republic of China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: XW ZF YZ. Performed the experiments: YZ XY LM ZF CB VV. Analyzed the data: YZ XY LM ZF. Wrote the paper: YZ LM VV ZF XW. Provided guidance on the whole study: ZF XW HW.