About the Authors

Thierry Olivry

Contributed equally to this work with: Thierry Olivry, Keith E. Linder, Ping Wang

tolivry@ncsu.edu

Affiliations Department of Clinical Sciences, College of Veterinary Medicine, North Carolina State University, Raleigh, North Carolina, United States of America, Center for Comparative Medicine and Translational Research, College of Veterinary Medicine, North Carolina State University, Raleigh, North Carolina, United States of America

Keith E. Linder

Contributed equally to this work with: Thierry Olivry, Keith E. Linder, Ping Wang

Affiliation Department of Population Health and Pathobiology, College of Veterinary Medicine, North Carolina State University, Raleigh, North Carolina, United States of America

Ping Wang

Contributed equally to this work with: Thierry Olivry, Keith E. Linder, Ping Wang

Affiliation Section of Medical Genetics, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Petra Bizikova

Affiliation Department of Clinical Sciences, College of Veterinary Medicine, North Carolina State University, Raleigh, North Carolina, United States of America

Joseph A. Bernstein

Affiliation Long Green Animal Dermatology Center, Baldwin, Maryland, United States of America

Stanley M. Dunston

Affiliation Department of Clinical Sciences, College of Veterinary Medicine, North Carolina State University, Raleigh, North Carolina, United States of America

Judy S. Paps

Affiliation Department of Clinical Sciences, College of Veterinary Medicine, North Carolina State University, Raleigh, North Carolina, United States of America

Margret L. Casal

Affiliation Section of Medical Genetics, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TO MC KL PW. Performed the experiments: PW TO SD JP PB KL MC. Analyzed the data: TO MC KL. Wrote the paper: TO KL MC PW. First recognized disease: JB. Helped characterize clinical phenotype: PB. Established pedigree: JP.